Ted Bennett said:Peter said:Quoted message said:Both pupil size and retinal sensitivity increase in response to low light levels. In bright light
the pupil size is around 3 mm and after some time in darkness it increases to about 5-7 mm. The
maximum size tends to decrease with age. I've never noticed any significant change in pupil size
based on focusing distance. Various drugs, either artificial or naturally produced by our bodies,
will also affect pupil size. The change in retinal sensitivity is over a much larger range than
that produced by changes in pupil size.
Ted replies:
Yes, that is why the variation in pupil size is more for focusing than for light control.
This is the opposite of what I've observed directly or read in other literature. It's clear that
pupil size varies rather dramatically with light level - I can readily observe this effect on my
own eyes while looking in a mirror and it is widely documented (e.g.
exploratorium.edupupil.htmlOpen ↗). And it is in a direction that makes sense for
dark adaptation by allowing more light to enter under dim light. OTOH, variations in pupil size
due to focussing distance appear to generally be very small.
It isn't the opposite of what you observe.
Your claim was that the variation in pupil size is primarily for aid in focussing and not for
adaptation to low light levels. [And I note you have yet to give us any reference to support
this claim.]
Yet if I focus on something at a short distance (which you claim will require a small pupil size)
under a low light level, then my pupil size is consderably larger than if I focus on infinity under
a normal light level. This is true for almost all people with some exceptions for people who lack
the normal pupil response to varying light levels. Therefore the pupil size is primarily determined
by light level rather than by focussing considerations. I.e. the observations are the opposite of
what I'd expect if your claim were true.
Quoted message said:How did you observe the effect of pupil size on image quality?
I have in the past observed the effect of looking through small apertures thereby simulating the
effect of a smaller pupil size. When observing at a distance where the normal accommodation of my
eye lens was still adequate to focus the effect of a reduced aperture is two-fold: 1) the brightness
is reduced as expected since less light is admitted, and 2) the image sharpness is decreased if the
aperture drops below about a mm due to increased diffraction effects. As long as the distance was
one where my lens still operated properly I never observed any overall image quality improvement by
artificially reducing the aperture.
OTOH, when observing at very close distances where the lens no longer has sufficient capability to
reach optimum focus (unfortunately this distance has been increasing for me lately) the reduced
apertures result in improved image sharpness while still reducing the brightness. This is the regime
where the 'pin-hole camera' effect mentioned by Carl becomes important, but note that it's ouside
the range of normal accommodation. However, even in this case the body does not have an effective
mechanism for reducing pupil size for an optimized image. Instead, this is the region where people
instinctively try to shut down the effective size of the eye with the eyelid in an attempt to
marginally improve the image sharpness (commonly referred to as "squinting" although the medical
community reserves this term for improper eye alignment which is something entirely different).
Quoted message said:
All right, let's try it this way: Variation in pupil size is more effective for focusing than for
light control.
The observed pupil variation over a size range of about 2 - 3 is quite effective at light control
since it results in a factor of 4 - 9 variation in the amount of light admitted.* OTOH, focussing is
controlled primarily by the lens and the eye is able to focus over almost as wide a range of
distances when fully dilated under dark conditions as it can when contracted under normal daylight
conditions although there is some reduction in close focussing ability at night. So no, I would not
agree with your new statement either.
I have tried to measure the effect on pupil size of changing the focus distance and was not
successful in seeing any consistent effect - AFAIK, the effect has been observed but is quite minor
compared with the size changes normally seen in response to light level changes.
I have already provided one reference on the relationship between pupil size changes and dark
adaptation and could easily provide more - do you have any quotable or easily-checked reference
that supports your claim that pupil size variations is *primarily* for focussing rather than for
light control? Or your new claim that pupil size is more effective at focussing than at controlling
light levels?
*Sure that's not much compared to the range provided by retinal sensitivity, but a factor of 5 times
as much light is nothing to ignore either. The effective light grasp of telescopes has also gone up
enormously as a result first of improved photo emulsions and more recently with better CCD
technology. But ask any astronomer if it wouldn't be nice to have a new space telescope with 5x the
area of HST or a land observatory with 2- 3x the diameter of Keck. For both telescopes and eyes, the
key to good low-light performance is to both improve the detector sensitivity and to increase the
aperture as much as possible. The dilated pupil makes the eye 5x as effective at dealing with low
light - show me the reference that says variation in pupil size improves the eyes focussing ability
by a factor of 5 or more.