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Absolute or just relative fitness?

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General fitness, health and nutrition
Published
5 July 2004
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5 July 2004
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John Edser
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  1. TT:- So: mentioning fitness is confusing, because of the
    many meanings of the word - and because the popular meanings
    lead to misunderstanding. A measure of absolute fitness
    would be more acceptable to me.

    JE:- Why will nobody here admit that Darwinism does contain
    an implicit absolute fitness?

    ______________________________________
    box 1 Darwinian fitness: The total number of fertile forms
    reproduced by each parent into one population.
    ______________________________________

    The net results of the Darwinian absolute fitness proposal
    above are:

    1) Darwinism can be experimentally refuted or uniquely
    verified testing Darwinian absolute fitness:
    __________________________________________
    box 2: Experimental test of Darwinian fitness. A natural
    population that experimentally holds the total number of
    fertile forms reproduced by each selectee within one
    population equal, must halt all natural selection within
    that population.
    __________________________________________

    2) Only a single level of selection exists within Darwinism
    as defined in box 1.

    3) Darwinian fitness cannot be selected to be reduced. Thus
    selectees are not forced to do anything except maximise
    Darwinian fitness. This singular nature of Darwinism
    mimics a teleological system.

    4) Just one case of organism fitness altruism documented
    within nature refutes the Darwinian single level of
    selection theory of natural selection.

    5) If the experiment in box 2 if verified then multi fitness
    levels are not verified so they can be validly removed
    using Occam's Razor

    Regards,

    John Edser Independent Researcher

    PO Box 266 Church Pt NSW 2105 Australia

    [email hidden]

  2. John Edser <[email hidden]> wrote or quoted:

    Quoted message said:


    TT:- So: mentioning fitness is confusing, because of the
    many meanings of the word - and because the popular
    meanings lead to misunderstanding. A measure of absolute
    fitness would be more acceptable to me.

    JE:- Why will nobody here admit that Darwinism does
    contain an implicit absolute fitness?

    ______________________________________
    box 1 Darwinian fitness: The total number of fertile forms
    reproduced by each parent into one population.
    ______________________________________

    The net results of the Darwinian absolute fitness proposal
    above are:

    FWIW, that isn't what I meant by "absolute fitness".

    If one is trying to refer to fitness as something maximised
    by evolution - or simply increasing - then this sort of
    thing won't do. Many organisms probably had about 2.6 kids
    millions of years ago as well - and this quantity shows
    relatively little sign of increasing with time.

    If you are trying to use fitness to measure progress in
    evolution, then you need a measure that really *does*
    increase over time.

    If today you replaced half of the world with its mesozoic
    equivalent - and then waited a few thousand years - I reckon
    most of the metazoic fauna and flora will have been wiped
    out in competition with modern forms - in a similar way to
    the way most marsupials don't stand up to competition with
    placental mammals.

    It would be a war of technology - the humans would
    obliterate the therapsids and archosaurs much as they are
    wiping out most of the modern large predators - by
    destroying or occupying their habitat - and by shooting
    them. The humans - and their ecosystem - would win because
    they have better technology on a range of fronts.

    It's perhaps a bit of a stretch to refer to something like
    "technological prowess" as a sort of fitness - but that's
    what I was talking about - not how many kids an organism
    has, is expected to have - or anything like that.
    --
    __________
    |im |yler timtyler.orgtimtyler.org [email hidden] Remove
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  3. "John Edser" <[email hidden]> wrote

    Quoted message said:

    Darwinian fitness: The total number of fertile forms
    reproduced by each parent into one population.

    Do you mean

    1. the number of offspring with the potential to reproduce
    if they live long enough (noting that most neonates are
    infertile - eg mammmals)

    or

    2. the number that actually survive to the age at which they
    become fertile (which will be <= the actual number
    produced)?

    ====================================================

    Do you mean

    3. "reproduced by each parent into one population." in a
    single mating

    or

    4. "reproduced by each parent into one population."
    throughout the lifetime of the organism?

  4. Tim Tyler <[email hidden]> wrote in message news:<[email hidden]>...

    Quoted message said:

    If one is trying to refer to fitness as something
    maximised by evolution - or simply increasing - then this
    sort of thing won't do.

    The appropriate maximand is something which might be called
    'relative fitness'. This obviously doesn't increase over
    time, since the expectation of relative fitness at all
    times is 1.

    Quoted message said:

    If you are trying to use fitness to measure progress in
    evolution, then you need a measure that really *does*
    increase over time.

    And there is no reason to call it "fitness".

    Wouldn't the sensible approach be to define "progress"
    first?

  5. Name And Address Supplied <[email hidden]> wrote or quoted:

    Quoted message said:

    Tim Tyler <[email hidden]> wrote in message
    news:<[email hidden]>...

    [number of kids metric?]

    Quoted message said:
    Quoted message said:

    If one is trying to refer to fitness as something
    maximised by evolution - or simply increasing - then
    this sort of thing won't do.

    The appropriate maximand is something which might be
    called 'relative fitness'. This obviously doesn't increase
    over time, since the expectation of relative fitness at
    all times is 1.

    That sort of relative fitness has some problems of its own.
    Who else do you include in the population? Only other
    members of the species?

    If so, that neglects competiton between species - and
    focusses on competiton between individuals - perhaps a
    rather distorted way to focus on nature.

    Quoted message said:
    Quoted message said:

    If you are trying to use fitness to measure progress in
    evolution, then you need a measure that really *does*
    increase over time.

    And there is no reason to call it "fitness".

    It seems like an appropriate term to me - if it wasn't
    already so overloaded. Today, "fitness" is a bad name for
    practically anything - due to the overloading problem.

    Quoted message said:

    Wouldn't the sensible approach be to define
    "progress" first?

    Progress means "advance" or "steady improvement".

    I did try to give some indication of what that might mean in
    the context of biology in my last post:

    ``If today you replaced half of the world with its mesozoic
    equivalent - and then waited a few thousand years - I reckon
    most of the metazoic fauna and flora will have been wiped
    out in competition with modern forms - in a similar way to
    the way most marsupials don't stand up to competition with
    placental mammals.''
    --
    __________
    |im |yler timtyler.orgtimtyler.org [email hidden] Remove
    lock to reply.

  6. If today you replaced half of the world with its mesozoic
    equivalent - and then waited a few thousand years - I reckon
    most of the metazoic fauna and flora will have been wiped
    out in competition with modern forms - in a similar way to
    the way most marsupials don't stand up to competition with
    placental mammals. It would be a war of technology - the
    humans would obliterate the therapsids and archosaurs much
    as they are wiping out most of the modern large predators -
    by destroying or occupying their habitat - and by shooting
    them. The humans - and their ecosystem - would win because
    they have better technology on a range of fronts. It's
    perhaps a bit of a stretch to refer to something like
    "technological prowess" as a sort of fitness - but that's
    what I was talking about - not how many kids an organism
    has, is expected to have - or anything like that.

    Tim Tyler

    ___________________________________

    Perhaps your mention of marsupials being outcompeted by
    humans was recently sparked by the following news story:

    Koalas face extinction, activists warn Australian group
    cites loss of habitat to development

    David Gray / Reuters file

    The future for these twin koala bears and their peers
    is bleak after an Australian group found that 30
    percent of 1,000 habitat sites surveyed no longer had
    koalas living there.

    SYDNEY - Koalas, Australia's iconic symbol, face extinction
    as rapid urbanization along the eastern seaboard destroys
    their fragile habitat, the Australian Koala Foundation is
    warning. The group has written to the government urging it
    to declare the koala a vulnerable species after a survey of
    1,000 koala habitats found 30 percent no longer had a koala
    in them and 60 percent had suffered widespread destruction.
    "I truly believe that in my lifetime the koala will become
    extinct unless we do something," said Deborah Tabarat,
    executive director of the foundation.

    Koalas are protected by law but the eucalyptus trees
    they call home and which provide their only source of
    food are not.

    There are about 100,000 koalas in Australia, down from an
    estimated seven to 10 million at the time of white
    settlement in 1788. In the 1920s, three million koalas were
    shot for their fur.

    Tabarat said the major problem facing koalas was that the
    majority of Australia's 20 million people and the majority
    of the koala population both call Australia's eastern states
    home. Eastern koalas extinct in 15 years? She said that with
    80 percent of Australia's east coast temperate forests
    destroyed and continued rapid urbanization, koalas along the
    eastern seaboard could be extinct in 15 years.

    "This animal is in serious trouble," said Tabarat. "In 15
    years you will not see a koala west of the divide," she
    said, referring to the Great Australian Divide, mountains
    that divide east coast Australia from its rural outback.
    Wild koalas only exist in four of Australia's six states:
    Queensland, New South Wales, Victoria and South Australia.

    The marsupial has no natural predator, but has been in
    decline for decades due to urban sprawl and from car
    accidents and dog attacks. More than 4,000 koalas are killed
    each year by dogs and cars, the foundation says.

    The most robust koala population on the Australian mainland
    exists in southeast Queensland and numbers about 10,000, but
    it too faces extinction in 15 years, said Tabarat. Southeast
    Queensland is experiencing the most rapid population growth
    of any part of Australia. Over the past eight years, 16,000
    koalas in the area died in accidents with cars or dog
    attacks and another 10,000 injured koalas probably died in
    the bush, said Tabarat. Fussy eaters

    Diminishing habitat has a greater affect on koalas than most
    animals. Koalas live in tall eucalyptus trees and low
    eucalyptus woodlands, but they are fussy eaters. There are
    about 600 species of eucalyptus in Australia, but koalas
    only eat about 120, with koalas in specific areas eating
    only four to six different types.

    An adult koala eats up to one kilogram of leaves each night.

    Like a pasture for sheep, a eucalyptus forest or woodland
    can only support a certain number of koalas, resulting in
    starving koalas in over-populated habitats or destroyed
    habitats. A koala population explosion on Australia's remote
    Kangaroo Island off the south coast has prompted calls for
    20,000 koalas to be shot to stop them destroying their
    habitat. The island has some 30,000 koalas struggling to
    survive. Koalas are also very social animals, living in
    stable societies that tend to remain in a small "home
    range," which means they require habitats large enough to
    support a healthy population and to allow for expansion by
    maturing young koalas.

    "People knock down all the scrub and leave a couple of trees
    and think koalas will be okay," said Tabarat. "We might be
    looking at koalas who are living happily in the bush but you
    might actually be looking at an extinct population," she
    said. "They haven't got any way of going out of their little
    home range, mating with someone then coming home pregnant.
    They just sit there, eke their time out, and then the bush
    will go silent."

    Copyright 2004 Reuters Limited.

    You state in part, "in a similar way to the way most
    marsupials don't stand up to competition with placental
    mammals. It would be a war of technology - the humans would
    obliterate the therapsids and archosaurs much as they are
    wiping out most of the modern large predators - by
    destroying or occupying their habitat - and by shooting
    them. The humans
    - and their ecosystem - would win because they have better
    technology on a range of fronts."

    Mr. Tyler, you are correct about humans technologically
    outcompeting marsupials on all fronts. Although I
    haven't checked the status of other marsupials certainly
    the koala is being outcompeted and there are a number of
    reasons for this.

    I live in one of the most highly "technologically" congested
    areas in the world and believe me quality of life
    considerations aren't taken into consideration...money is.
    It's interesting how you dichotimize between the ecosystem
    of marsupials and placental mammals. In the past many other
    animals became extinct or close to extinction because of
    humans shooting them for their fur, tusks, body parts, etc.
    In fact, many other animals and plants face extinction
    besides marsupials because of rampant human development and
    human's "technological prowess".

    I hesitate to call it "technological skill'" because in my
    view it is being misapplied. This is through no "fault" of
    our own as we are just following the dictates of Darwinian
    evolution and we happen to be the Alpha animal which can
    destroy anything. But this does potentially have
    consequences in the long term..this rampant development of
    our "technological prowess".

    Many have remarked on the destruction of the rain forests
    and the potentially medically and scientifically valuable
    specimens in them, not to mention the rain forests provide a
    certain percentage of life giving oxygen to our atmosphere.
    The more rampant development of our "technological prowess"
    without regards to the environment (our environment),
    demographics of population, service needs, water, potential
    pathologies which result from such conditions, etc, etc. we
    will indeed be not only negatively effecting the ecosystem
    of marsupials and other plants and animals but ourselves.

    In fact, I think we've already arrived there Mr. Tyler.
    There are over 6 billion people on earth and overpopulation
    is a serious threat. Now perhaps you think such concerns
    aren't warranted as eventually massive natural selection
    will kick in and wipe out a signifigant percentage of the
    human race. But if that happens it seems our "technological
    prowess" will have somewhat backfired on us and the
    consequences are unpredictable.

    Michael Ragland

    "It's uncertain whether intelligence has any long term
    survival value." Stephen Hawking

  7. Michael Ragland <[email hidden]> wrote or quoted:

    Quoted message said:

    [...] we will indeed be not only negatively effecting the
    ecosystem of marsupials and other plants and animals but
    ourselves.

    In fact, I think we've already arrived there Mr. Tyler.
    There are over 6 billion people on earth and
    overpopulation is a serious threat. Now perhaps you think
    such concerns aren't warranted as eventually massive
    natural selection will kick in and wipe out a signifigant
    percentage of the human race. But if that happens it seems
    our "technological prowess" will have somewhat backfired
    on us and the consequences are unpredictable.

    I don't forsee much in the way of population crashes.

    I found an interesting graph relating to human population
    growth through 2150 recently:

    n_Growth.htm They say:

    ``In 2000, the world had 6.1 billion human inhabitants. This
    number could rise to more than 9 billion in the next 50
    years. For the last 50 years, world population multiplied
    more rapidly than ever before, and more rapidly than it will
    ever grow in the future.''

    Their projections seem fairly realistic to me - though
    perhaps they are on the pessimistic side - surely the world
    population will reach 11 billion *well* before 2150.
    --
    __________
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    lock to reply.

  8. Tim Tyler <[email hidden]> wrote or quoted:

    Quoted message said:

    I don't forsee much in the way of population crashes.

    I found an interesting graph relating to human population
    growth through 2150 recently:

    n_Growth.htm

    The s.b.e long URL bug strikes :-|

    Try:

    tinyurl.comsns7

    ...which leads to the page I originally referenced.
    --
    __________
    |im |yler timtyler.orgtimtyler.org [email hidden] Remove
    lock to reply.

  9. Guy Hoelzer said:
    Quoted message said:
    Quoted message said:

    EK:- Do you mean
    1. "reproduced by each parent into one population." in
    a single mating or
    2. "reproduced by each parent into one population."
    throughout the lifetime of the organism?

    Quoted message said:
    Quoted message said:

    JE:- Obviously, you would have to exclude 1. The total
    number of fertile forms reproduced into the same
    population by one parent may not be completed until well
    after that parent has died. Plants can leave infertile
    embryos (seeds) dormant for centuries. They may end up
    becoming fertile forms many years after the death of the
    parent plant. The total number of fertile forms
    reproduced by a Darwinian selectee over its lifetime
    just is a good approximation.

    Quoted message said:

    GH An approximation? This seems like a major departure
    from your long standing insistence that this is exactly
    the Darwinian fitness.

    JE:- I defined Darwinian fitness in box 1. Logically this
    definition allows for a parent to die before all of its
    infertile reproductives reach fertile adulthood. Mostly
    this is not the case so mostly, the total number of fertile
    forms raised to fertile adulthood within the same
    population over just a parents lifespan is only a good
    approximation. The totals remains FINITE within both the
    definition and its approximation so both remain testable
    suppositions of nature.

    Quoted message said:

    GH:- I have consistently argued that there can never be an
    exact definition of fitness, because I see fitness as
    merely a heuristically useful concept, demanding a variety
    of operational definitions in its application, rather than
    something that actually exists, demanding a single
    accurate definition.

    JE:- I have consistently argued that fitness is an
    OBJECTIVE ABSOLUTE ASSUMPTION OF NATURE upon which all
    evolutionary theory is based and upon which the
    refutability and verification of ANY theory of evolution
    depends. I have unambiguously defined fitness and given an
    experimental confirmation of that definition that only
    requires a single level of selection. You have denied that
    selection can be halted within any natural population using
    the exp. provided only because for you, multi levels of
    selection exist in nature. You will not even consider a
    single unit alternative yet you offer no refutable position
    on this matter. In short, you wish to substitute just your
    hand waving exercise of only heuristic multi levels of
    fitness for a more simple, objective, testable and a more
    simple single level theory of fitness. What you keep on
    offering here is just, bad science.

    Quoted message said:

    snip<

    Respectfully,

    John Edser Independent Researcher

    PO Box 266 Church Pt NSW 2105 Australia

    [email hidden]

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